Isopod Care Guide Part 7: Breeding Isopod Morphs
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- Опубликовано: 13 сен 2024
- Isopods are some of the best pet invertebrates! In this video, I discuss how to breed isopods selectively to create morphs. I describe my experience producing orange Dalmatian Porcellio scaber starting with orange Porcellio scaber and Dalmatian Porcellio scaber isopods.
Check out these LINKS to Isopod foods and supplies below. As an Amazon Affiliate, I earn from qualifying purchases.
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Wally at Supreme Gecko makes a great Isopod food: www.supremegeck...
superior shipping supplies.com makes the smaller isopod enclosures I show in the video.
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Isopods will munch on dried River shrimp as well!
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Isopod Substrate ingredients:
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I am a participant in the Amazon Services LLC Associates Program, an affiliate advertising program designed to provide a means for sites to earn advertising fees by advertising and linking to amazon.com. In other words, I earn a small commission when you use that affiliate link, at no cost to you.
I stumbled across one of your videos about a week ago and I feel as though I've entered a totally new and fascinating world. As a result, I have my first isopods on order and can't wait for their arrival. I am a veteran of invertebrate culture and over the years have kept phasmids, snails, millipedes, cockroaches, centipedes... but I'd always overlooked the lowly isopod. What have I been missing?! So, thanks, Rus, for inspiring me!! BTW, I live in the south of Spain and as Spain is a hotspot for isopods, it will be interesting to see what I encounter locally.
Welcome to the world of isopods! I’d love to hear which species you see in your area.
Been mostly a tarantula guy for 10 years at this point. Now I'm 10 varieties of isopods deep for both CUC and display, a colony of blue death feigning beetles and pining after more and more arthropods because of russes videos
This is probably the coolest hobby because you can do it without any money.
For about a year I’ve been slowly working on a “golden” morph with originally only grey and yellow-spotted isopods. Right now I have a good population of butter-colored pollies :) I’m now working on rose colored ones and calicos! All from wild caught isopods!
Do you have any idea what species you have?
woah that’s amazing! do you have any tips on doing this?
I believe they are Armadillidium vulgare!
Lots of patience and basically weeding out the ones that didn’t take after the butter-colored morph. I accidentally forgot to do this for a while and many of my golden’s babies ended up grey again :’) there’s more to maintaining the genetic pool, but I’m a newbie, so I think luck was just on my side 😂
@@justsomerandomperson761 interesting! sounds like a fun project
This is an absolutely brilliant video and one that I was definitely waiting for!
Early I am working with a magical dairy cow gene because they pop up in every single bin I have! These guys are like mites, they just show up everywhere! They must be masters of escape.
I think there is indeed an ‘Apparate’ gene in several species of isopods. P. pruinosus, T. tomentosa, C. convexus...
I found some orange scabers local to me last year and I’m working with those to try and get them to produce all orange. Also getting some calico looking ones in the mix.
That’s great! Good luck with your breeding project 👍🏽
My first Isopod Culture I made was actually wild caught P Laevis Cali Mix from my yard. (I'm still very new to isopod & Invert keeping in general.)
I've had been actually trying to take out the individuals of wild types in the culture,while only keeping the morphed individuals. After several months of taking out babies and subadults that were wildtype, I've noticed much more of the offspring kept that Cali-Mix traits. I get tons of cream and pied individuals. Even a very select few that have mostly dark grey with little white and orange splotches around the skirts, for every 10-15 of those morphed individuals I may get a single wildtype.
I may breed them with dairy cows eventually to see what could happen. Once I get a well established population made.
Thank you so much for the informative video!
Interesting! I’d love to know how the genetics of that morph work...
I Like Hearing about your selectively bread wild strains
Thanks for the video! I'm a raw beginner to the hobby so I got a lot of useful information.
There are no isopod breeders near me, so I got my first colony by collecting wild specimens from my garden a few months ago. I have no idea of their species. I live in Australia, but since I got the isopods from my garden, I'm guessing they're not native. I recently moved them to a larger enclosure and got a pleasant surprise. There were a number of individuals that were pure white! I put them in an enclosure of their own in the hope I'd breed more, but from what you're saying there's probably a much lower chance of that happening than I'd thought. Still, I'm enjoying myself a lot.
I also noticed I have a few orange individuals, so I may just have a consolation prize. :D
wow, those isopods that you found must be carrying lots of interesting genes, telling by their offspring! maybe you can become your regions first isopod breeder;)
Explains why some of my Isopods in my community tank are mottled deep grey and orange, similar to the 'Lava' type.
I am just enthralled in the genetics of Isopods. I've kept many plants and crossed different colors and species throughout the years (Even ended up with a Hedera Helix plant crossing with my Japonica, but none of the young survived for longer than two years).
I just wish there was a cheat sheet for what to cross and what to look for. I have so many questions, but I don't know how to ask them.
Edit: Fatsia Japonica*
I think that is how they started the hybrid called Fatshedera lizei (Ivy tree) - infact I have some form of Ivy crossed with fatsia japonica myself I found for sale in a florist stall year's ago but it's always remained a rather feeble little specimen unfortunately. All its syblings I bought at the same time died some years ago now. I also keep tarantulas & centipedes & a small number of isopods along with various other inverts - it's an illness, Im never satisfied with what I have! Lol
these videos bring me so much joy, my day is so much better when I see them in my subscriptions
I really appreciate the knowledge you share. I just found the first mancae in my culture of speckled armadillium vulgare that I separated from my colony of wild caught ones. I can't wait to see what patterns they'll have when they grow out a bit.
Great video as usual! Perfect timing as well, I've been wanting to get into breeding isopods.
Another outstanding video Rus. You cover all the points so well. Your knowledge is amazing. Had this video topic on my list for 2 weeks from now :) Glad you beat me to it as you did a fantastic job! I get this question almost daily and will start pointing people to this video.
This is so fascinating. Thank you for sharing all this info in such detail. I’m about to receive my first isopods in a few days, and didn’t have any desires to breed a colony or breed morphs, but your video has got me thinking twice.
I'm kind of anxious in several ways about starting down the breeding aspect. I feel you on that!
And similarly, the more of Rus I watch, the more I'm willing to do more!
@@MIsopods Good luck! I hope you have a fruitful journey. Our isos have been with us since October and so far, no successful breeding which is better for me at this point. We’ve had a couple die though and many failed attempts by the littlest ones to mate with bigger ones. 😂
@@lorraelena well I wish you more and better luck! I've had similar experiences with all of my starter colonies. Very slow to establish themselves.
I mentioned to Rus in another video, that thanks to his videos, it helped curb my anxiety about a lot of that. I think I mentioned to him about how, his and other channels show population explosions, and that is not generally a realistic expectation.
As long as I know things go slow, I'm ok with it.
keep me posted!
I saw this because I ended up with an orange-brown garden roly poly and was wanting to breed more because they are so cute. This helped a lot, thank you!
Great stuff Rus. I thought I wouldn't ever want to purposely breed any animal but that was coming from hanging around in the small mammal and bird communities. Isopods however seem a much more acceptable "risk"/challenge, and are fascinating.
I’m late but I have noticed a newish color morph in my orange Dalmatians. It’s this reddish orange brown color. Been calling them Autumns. They are a soild color too. So far there is four. One adult and three babies. I really like this color so I am going to remove them and put them in their own bin later tonight
They sound pretty! I’d like to see them
Today I found what appears to be a red Dalmatian or maybe Calico Vulgare. Someone pointed me to this video and I hope it will be helpful in my pursuit of isolating this beautiful trait!
Loved the video. I'm new to isopods so I don't have any of my own projects but I am interested in what colors my new P. Scaber lotto mixes are going to be. A few of the first ones are starting to get their color. I also have a bunch of new ones that I know were from a beautiful red calico in the culture. Can't wait to see them grow and discover what they are.
That is one of the joys of lottery mixes, definitely!
I'd love to see them! It is so interesting what colors and morphs can pop up!
@@RealBradMiller I ordered a 10 count and got 13. There were 3 visibly wild types (possibly male calicos) 1 calico, 1 red calico, 3 oranges, 1 dalmatian, 1 calico dalmatian, 3 orange dalmatians. There is lots of potential for suprises :)
@@RisingFromTheAshes97 That's awesome! Are you adding them to an existing colony or are they on their own?
If this is your first time with Isopods, I recommend using a red LED flashlight to view them at night, that's when you really see some activity!
@@RealBradMiller This is my first time with them. I have some wild caught A. vulgare's and I bought some A. vulgare orange vigors, P. dilatatus, the P. Scabres and Oniscus asellus Mardi Gras Dalmatian. I figured i'd get 10 counts of each and see how they did. So far they are all doing pretty good. My wild A.vulgares and the scabers have both bred.
I've just started keeping isopods. My one colony as of now is all wild type A. vulgare collected from around my house (we get a *lot* of them inside during the warmer months). I'll be keeping an eye out for neat traits as they get established.
Your videos have done a lot to help me get started! Thanks for putting all this info out here, and best of luck on your projects!
Great video.
I'm working with zebras at the moment and have isolated some cream/white specimens and now have around 10 small unmated ones.
Started off with 2 semi creamy ones with white antennae/legs and some other lighter colour zebras.
Im hopefully going to get mainly lighter ones in my third generation.
I deeply respect the amount and quality of knowledge youve shared! Thank you! This is so fascinating. Do you know any websites/documents/books that give at least an outline of what kind of genes/characteristics are recessive and dominant in isopods? I'd love to know more.
I am very happy to know you found this useful! I am not aware of a resource with that information (though there may be one.) Most mutations seem to be simple recessive traits, but many calico forms of P. scaber are sex-linked. Porcellio scaber ‘lava’ does not seem to be simple recessive.
good stuff
This is so informative! Great job, fantastic reporting!🌻🌼🐝 Keep it up 🙌
GREAT video Rus... I've got a high yellow A. vulgare breeding program that's around 9 months in... I've now started to selectively take out the more noticeably colored critters and isolating them in another enclosure...
Awesome video rus
Thanks Frank!
Sometimes YT randomly recommending channels is spot on, I have loved isopods for quite a while and years ago tried breeding them for the first time as a kid. That shot at the end was gonna be my prize line after i got ahold of a single armadillium wild caught that had an orange back and two or three with white instead, now i think i sorta understand why i got a dozen orange and yellow and only three or four white offspring in the first generation and maybe a single white marked one in the second. Just recently got new wild caught ones, think they're scabers if i am remembering right, i got lucky and found a spot nearby with a mix of wild grey, some with a slightly stony texture up close and less solid grey and some powder blues, a few of which are paler. I only took a few immatures of both of course since i want the wild population to keep breeding but i definitely separated the shiny shelled powders from the greys and may actually split them up into three or four colonies after the powders mature and start reproducing, since I'm curious if separating the paler specimens of that particular group could eventually yeld a silver morph, but if not, at least that group has nice color either way. Thanks for the video and I'll def keep a closer eye on what morphs interact with what from now on in case i figure out something I really like looking at, though at least for now I'm just keeping them for personal enjoyment and honestly don't know what else I could do with them. I have only barely gotten into looking into what custom morphs are already on the market so I'm unlikely to breed anything new myself, since I haven't looked around enough to figure out what might not be already out there
I’ve been waiting for this one, boyes. Let’s go!
Russ - you legit inspired me! I came to your channel earlier this year over I think neocaridina, and I now have 7 species of iso's. The color morphs of neo's interested me but so much work had already been done and line breed culling seemed way too complicated. The second video of yours I watched was your dairy cow + orange. That inspired my inner child "mad scientist" of creating some new life form. I emailed you a while back about trying dairy cow with cali mix as I haven't seen those attempted yet. I don't have much hope for success, however, I upgraded my dairy cow enclosure twice, and the cali mix got the old enclosure twice. The first has a tooon of mancae from both and NO adults (who were moved 2-3 weeks ago) - I think this is going to be my best chance. They are all roughly the same age probably coming up on a month old now so pretty sure none are sexually mature yet.
This is likely the fifth time I've watched this video. I'm currently working on recreating orange cream also. I just started my lava colony, today (as well as expansus) from smugbug. Once I grow my lava colony out I want to play around. I still don't understand the co-dom and asked Luara at smugbug if she could explain it better. Your explanation helped a lot, as I would have never thought to cross it back with a wild type. I'd love to see or hear what you have done with lava's since this video is a few months old - I'd love some ideas on what to cross them with. Orange dalmatian was my first thought. The most interesting and longest project I'm just starting is trying to line breed blue dairy cows. Calling all dairy cow owners, I'm interested in collecting any cows that throw a good shade of blue. I don't mean wild-type silver/grey-blue, but blue blue. I know line-breeding is going to require starting with more of a wild blue and take MANY generations to get a blue, but that is now my life goal!! If I ever get it I'm naming them Buggalo, in honor of futurama that dairy cows remind me so much of.
I'm not sure how you keep your notes on your breeding experiments, but I would love to hear more about any of your successes and failures to save me at least a little struggle!!
The next species I hope hits sellers fairly soon is Merulanella sp. “Tricolor” and Merulanella sp. ember bee. I know everyone buying them right now are seeing $$ signs, I just want those gorgeous animals in my collection.
I love too see comments like yours...knowing that I have helped inspire you to delve into projects like these. As far as crossing the Lavas with another morph, my suggestion would be either pied or orange pied. Hopefully this would lead to true tricolored individuals with sharply defined patches of color.
If I were to start another laevis cross attempt, it would like be Cali Mix with orange, as I believe this would be a compatible cross.
I love the idea of a blue P. laevis...hopefully we can eventually get one not caused by iridovirus, or at least by a strain that is not harmful, like the strain the causes a blue color in some crayfish.
I hope this helps, and keep it up!
Awesome video mate as always :)
I was wondering how to create different morphs and looking for proper info online, but your video clearly answered all my questions... Thanks! :)
Thanks for the knowledge!
Working on my own nightshade armadillidium vulgare. Also Working on powder oreo crossing with powder orange
I’ve been setting up an attempt at a breeding project for my a. Vulgare isopods- I think it’s similar to your nightgold line, since I’m focusing on the yellow spots of them :D
Thank you Russ, this was very interesting.
I have vulgare wild types right now. I'm hoping that as they grow they express different traits at the colony matures. I have about 12 wild caught individuals. We'll see how it goes.
I finally found some today and a few have the gold fleck. ( I wonder if this is common)
Having a few good flecks is quite common in females. Having most of the carapace marked with gold is quite uncommon.
great video Rus!
Fascinating story!
hello great video information Russ
Hi. Great Information. The Armadillidium vulgare you are showing near the End of the Video look pretty similar to the Morphs in my A. frontetriangulum-Colony. And today i spotted a second one. I will try out your tips and let you know, if it works. Thank you very much
Thank you for your great info! I have some A. maculatum that are the "chocolate" colour, mixed in with my regular black and white ones. I'm trying to breed some "Chocolate Zebras" now, after separating the ones with the desirable colour. Most of the new babies have the black and white colour, but there appears to be one chocolate baby! I will keep their little group together and see what happens. I only started this a couple months ago.
Do you think it’s possible to crossbreed Armadillidum Werneri with klugii ?
I’ve been interested in isopods for ages but just now thinking about getting some to start my own hobby colonies and see if I can get any cool morphs :)
Welcome to the hobby!
Wish for a new video. Why do expressed Recessive types produce wild form offspring? More about genetics
I've got an Orange Piebald Scaber line going and the last generation is growing up to hopefully prove them. Although it had a recessive whiteout gene mixed in that cause some to look like Yellow Snow and one was pied, so my guess is triple recessive; pied + whiteout + orange.
Fascinating how different genes can combine, especially since when whiteout is typically expressed it seems to remove all pigment.
@@Aquarimax Tyrosine can be in a T+ morph, or maybe a hidden yellowing gene for the orange + whiteout removed melanin but keeping Tyrosine. Then piebalding removed them both.
I brought in just two local wild ones, thinking vtgey didn't like each other, however months later, babies in ky terrarium
What's interesting about the "Ukraine Pied" Cylisticus is that they look like a typical dalmatian morph more than anything. Whereas the "US Pieds" that never bred true had much less small grey speckling, the grey and white blotches were bigger and more clumpy, and some individuals were just pure white if I recall correctly, with others being almost pure grey, but with one or two white spots. None were quite as evenly mottled as we see in the "Ukraine Pied" stock.
I keep my Dairy Cows and my Giant Canyons together. They frequently try to mate but I haven't seen anything to indicate they were successful. They both are breeding very well with their own morphs though.
Cool! How long have you had them together?
@@Aquarimax I've switched enclosures on them a few times but they have been rooming together for 6-7 months in total
Ok, so for the moment, I'm trying to figure out how to keep isopods in a very small environment, while being able to breed them, and by small i mean a container of the size of a human fist ✊ (a male human)
I just stumbled upon this video while trying to figure out this stuff, and now i just need to know what isopods need for general survival
Definitely check out my playlist on isopod care! There are some species that could do fine in a small environment like that .
I found two that's half white and half dark Gray
Hey Rus. i know this is an old video but I was wondering in situation like this where you are separating out a small handful of isopods for selective breeding, how big of a container would you suggest housing them in.
Even when starting a new colony I worry about putting them in something smaller than a 6 quart because it’s a little harder to control humidity gradients but also putting them in something like a 6 quart seems like it’d be possible it’d be too big and they would have trouble finding each other to breed.
I’d love to hear your thoughts and advice if you see this and get a chance. Thank you and keep up the good work !!
A few months ago I found some armadillidium vulgare in my garden that looked very different then the other pill bugs I put them in a container and now the pill bugs produced babies
Im about to go wild isopod hunting in England it will be fun to see what happens in the coming year. Is it possible to line breed from a wild caught colony after its established or are wild ones unlikely to mutate ?
I've found various orange Armadillium vulgare around my backyard in the wild.
I've collected them and now I just finished their enclosure.
I may order some Armadillium vulgare orange to breed with them.
I also have a few very large Armadillium vulgare gold .
I only have wild caught large individuals with that patterns.
I've never found any small ones or anything like that.
I'm not sure if they look different as a juvenile or if the pattern shows up much later.
I put all of my orange and light colored isopods in a new enclosure.
l remember soooo many armadilliums around when l was growing up in the city.
Now, all l don't seem to be porcellios or rathkii.
Awesome.
Third attempt to get these delivered, it's 9° outside. 3 attempts all DOA
Oh no! I generally take a shipping break for the worst of the winter. I hope you are able to get a viable order!
Ive been keeping A.Vulgare and P.Scaber colonies that are descendants of specimens I caught in the wild and I keep all my favorite looking A.Vulgare specimens in one bin together. recently I spotted a single bright orange young one among the masses and it sent me into such a frenzy!!!
I recently bought a small count of "punta cana" morph A.Vulgare and I am curious to know what adding them to the colony will do to the gene pool and if you have any insight for me in that regard.
On a similar note, Ive done the same with my P.Scaber colony and even though they are all visually wild type, theyve produced a few offspring that are calico, white with orange speckles, and a single cream colored one. What would happen if I were to introduce about 5 "koi" morphed P.Scabers to the mix? I'm not familliar with the genes I'm working with.
Sounds like some fun projects you are working on! Typically, the best way to work with a specific trait is to isolate any individuals with that trait along with a few young wild types, that way in a couple of generations you have a chance of finding other individuals with the same trait, without being confused by the interactions with other traits.
You could add ‘Punta Cana’ with your Armadillidium vulgare projects, for example, but then it would be very difficult
to attribute the potentially very variable phenotypes possible offspring of subsequent generations to any of the other stock y out introduced.
If you introduced ’Koi’ stock to your P. scaber projects, the first generation would likely
Look like wild types, (if they are all recessive traits) and then you would likely need to get another generation from those offspring before you could see how the genes interacted, but it would be difficult to know which genes do what. So, to get an idea of how /if each gene is heritable, I suggest focusing on the stock you have along with wild types. 👍🏼
I actualy found some wild Lava Isopods in my garden. 2 of them were like this. I wonder why?
They might be Calico, that mutayseems more common in wild stock than Lava
Whooop
I am wondering, is it possible to breed say, a lava isopod that has a codominate trait with a Dalmatian that is a recessive trait?
Have you tried breeding with like orange males and cow females only? Or other way around? So they couldnt possibly mate with their own color.
So cool. How hard is it to start doing this with no prior isopod experience?
I would say if you keep some isopods for a couple of months successfully, you can go ahead and try it.
@@Aquarimax my man thanks for the info
@@joshsaunders6392 I'm not very smart, and even I can do it! Haha.
Take some time and do some research, and I think you will be fine.
@@MIsopods lmao ok thanks any recommendations on where to learn the info?
@@joshsaunders6392 RUclips keeps removing my replies...
Can these affect asthma ? I have asthma and dubia roaches affect them but what about these? Do they make you sneeze or make it hard for you to breath
I am afraid I am not qualified to answer that question, you may want to speak with a doctor about that.
Are working on breeding across of powder orange and dairy cow
they are different species so it won't work
Why do you think Dairy Cows are more successful than other Isopods? And how do you pick up young isopods without crushing them?
I had a handful of wild type vulgare pop up with some having white on their bodies
Awesome! Worth working with!
I also have a question about texture mutations...
In my A.Vulgare colony, after a few generations of undisturbed breeding, I selected all the babies I could find with any sort of eye-catching uniqueness to them, and put them all in a separate bin. After a few months had passed and most of the young had become big enough to properly appraise, I noticed two individuals that looked very similar to each other but nothing like the rest of the colony. Their shell segments were lifted and flared out a little bit from the head, their body was a very light gray with light yellow markings, and the shell had a noticable rippled/bumpy texture. The only time Ive seen this shape and texture, is when I'm looking at a different armadillidium species entiely! (granulatum I believe) Any insight?
I have heard of a few of these texture mutations…in Porcellio laevis and Armadillo officinalis, I believe. Since you have two, you have a decent chance of isolating the trait if you keep them with a few wild types, preferably from the same stock, and seeing what happens in a few generations. I would keep a close eye on them though. Some texture changes could result in molting issues or other problems, so keep an eye out for that
OK so what if I took my Orange Dalmatians and added some oranges just like you did here? Would I be able to produce orange Dalmatians with more orange to them? Cuz mine don't have all that much orange to their patterns.
Unfortunately, it doesn’t work that way. You would likely fry solid oranges in the F1s, and some Orange Dalmatians in the F2s, but it wouldn’t necessarily increase the orange expression. What you might want to do is work with Orange Koi or Orange Piebald, they tend to have more orange expression.
Awesome. Thanks for that info.
I caught 50-ish Armadillidium Vulgare a few evenings ago. Upon further inspection, I noticed that some of the brown/caramel colored Vulgare have these really neat red markings on their tail section. Some look like paint brush streaks and some look like a red skirt that just goes around the tail. Is this a common marking? Does anyone have any experience or information on this trait? I'll try to get some photos also.
That is not something I see often . Would like to see some pics!
Porcellio Laevis Dairy Cow and P.L. Orange can't produce offspring? Bummer, I just bought some P.L. Orange intending to put them in my D.C.s' bin and get some orange cows, lol.
Can they at least live together harmoniously since they are both P.L.? As long as there is enough food I mean.
It Appears that they are no longer considered the same species, BUT I have Milkback and Orange living together, they’ve been together a few years, and they are doing well 👍🏼
I have powder blues and some orange among them. Do they cross?
Yes, powder orange and powder blues will readily reproduce together. Orange is recessive. 👍🏼
@@Aquarimax thx for the response!
I've accidentally become obsessed lol
It happens! 😁👍
So if you breed a porcellio scaber orange with a dalmation you get a porcellio scaber orange dalmation. But what do you think would happen if you use a porcellio scaber lava instead of orange
Good question! Lava seems to be co-dominant, so you might get a low_expression lava, but I hope people test it out soon!
@@Aquarimax i am defenitly going to try this then!
What is a “wild-type”? What is a calico?
A wild type is what the species looks like in the wild (gray for many species) and a Calico has random marking of 2 or more colors. 👍🏽
Does anyone know if Porcellio laevis milkback and porcellio laevis dairy cow will breed together?
I believe they will, but I haven’t tried…I feel like they are similar enough that the offspring could be inconclusive, and if the offspring get out into the hobby, they could co BFF use things further.
Love that shirt 😏
Thank you!! 😊👍
I have one question about keeping and breeding Isopods .... ahh .... WHY??? lol
The same reason people keep spiders or fish. It's a fun hobby.
Why does it look like he is wearing a sleeve of hair.
Are you freeing to my arms?
I have milkbacks and dairy cows that I’m going to try and breed once their populations are large enough to start a third tub, I’m not sure what will happen but it will be cool to see what happens
It has been done, but I am not sure I should spoil the surprise 😁
hello would this hold true with roaches ? do you know anything about the golden dubia roaches , or the john forest amber / yellow dubia roach breeding program
There are some general similarities with breeding roaches as well, from what I understand, though I have never bred roaches. I have heard about the golden Dubia projects, on Kyle’s livestreams (Roach Crossing)